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Alberto Esquivel, Arne Lesterhuis, José Luis Cartes, Rebecca Zarza and Hugo del Castillo and are used with their permission.
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BIBLIDINIAE - TRIBE BIBLIDINI - BIBLIDS
A large and varied group of small to large Nymphalid butterflies which includes distinctive groups such as the 88s, the Clickers and Sailors. They occasionally are grouped under the common name "Tropical Brushfoots". Amongst the most charismatic species are the "Clickers", capable of producing a crackling sound like electricity with the wings (see Gallery 4 for photos and to hear the noise). They are common inhabitants of forest undergrowth and gardens where they perch with open wings on tree-trunks and walls. Butterflies in the genera Callicore and Paulogramma are easily mistaken for the true 88 butterflies of the genus Diaethra, but note the yellow rather than white base colour of the underwing.
Click the photo to enlarge the image, click the links for the FAUNA Paraguay Handbook accounts.
FIGURE 1a

FIGURE 1a
FIGURE 1b

FIGURE 1b
FIGURE 1c

FIGURE 1c
FIGURE 1d

FIGURE 1d
FIGURE 1e

FIGURE 1e
FIGURE 1f

FIGURE 1f
FIGURE 1g

FIGURE 1g
FIGURE 1h

FIGURE 1h
Gallery 1 Biblis, Callicore, Catonephele and Colobura
FIGURE 1a - Red Rim Biblis hyperia - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 1b - Little Callicore Callicore hydaspes - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 1c - Little Callicore Callicore hydaspes pair mating - Hotel Tirol, Departamento Itapúa (Regis Nossent July 2005).
FIGURE 1d - Blue-bordered Callicore Callicore pygas - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 1e - Zigzag Callicore Callicore sorana - PROCOSARA, PN San Rafael (upper Paul Smith March 2007, under Martijn Brouwer May 2007 - ECOSARA Biodiversity Database).
FIGURE 1f - Stoplight Catone Catonephele numilia male - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 1g - Stoplight Catone Catonephele numilia female fulva form - Hotel Tirol, Departamento Itapúa (Regis Nossent July 2005).
FIGURE 1h - Zebra Butterfly Colobura dirce - PROCOSARA, PN San Rafael (Nick Simms February 2008 - ECOSARA Biodiversity Database).
FIGURE 2a

FIGURE 2a
FIGURE 2b

FIGURE 2b
FIGURE 2c

FIGURE 2c
FIGURE 2d

FIGURE 2d
FIGURE 2e

FIGURE 2e
FIGURE 2f

FIGURE 2f
FIGURE 2g

FIGURE 2g
FIGURE 2h

FIGURE 2h
FIGURE 2i

FIGURE 2i
FIGURE 2j

FIGURE 2j
FIGURE 2k

FIGURE 2k
FIGURE 2l

FIGURE 2l
FIGURE 2m

FIGURE 2m
Gallery 2 Diaethria and Dynamine
FIGURE 2a - Number Eighty Diaethria candrena - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 2b - Number Eighty Diaethria candrena - Hotel Tirol, Departamento Itapúa (Regis Nossent July 2005).
FIGURE 2c - Number Eighty Diaethria candrena - Hotel Tirol, Departamento Itapúa (Regis Nossent July 2005).
FIGURE 2d - Number Eighty-Eight Diaethria clymena - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 2e - Edged Sailor Dynamine agacles - Encarnación, Departamento Itapúa (Paul Smith June 2007 - ECOSARA Biodiversity Database).
FIGURE 2f - Edged Sailor Dynamine agacles - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 2g - Small Eyed Sailor Dynamine artemesia male - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIIGURE 2h - Small Eyed Sailor Dynamine artemesia female - PROCOSARA, PN San Rafael (Paul Smith June 2007 - ECOSARA Biodiversity Database).
FIGURE 2i - Lined Sailor Dynamine athemon - PROCOSARA, PN San Rafael (Paul Smith November 2007 - ECOSARA Biodiversity Database).
FIGURE 2j - Whitespot Sailor Dynamine coenus - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 2k - Whitespot Sailor Dynamine coenus - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 2l - Four-spotted Sailor Dynamine postverta  - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 2m - Blind Sailor Dynamine tithia - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 3a

FIGURE 3a
FIGURE 3b

FIGURE 3b
FIGURE 3c

FIGURE 3c
FIGURE 3d

FIGURE 3d
FIGURE 3e

FIGURE 3e
FIGURE 3f

FIGURE 3f
Gallery 3 Ectima, Epiphile, Eunica and Haematera
FIGURE 3a - Ectima thecla - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 3b - Epiphile hubneri - PROCOSARA, PN San Rafael (Paul Smith November 2007 - ECOSARA Biodiversity Database).
FIGURE 3c - Ivory Eunica eburnia - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 3d - Ivory Eunica eburnia underwing in cryptic pose - Encarnación, Departamento Itapúa (Paul Smith June 2007).
FIGURE 3e - Purplewing Eunica tatila male - PROCOSARA, PN San Rafael (Paul Smith September 2007 - ECOSARA Biodiversity Database).
FIGURE 3f - Blaze Haematera pyrame - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 5a

FIGURE 5a
FIGURE 5b

FIGURE 5b
FIGURE 5c

FIGURE 5c
FIGURE 5d

FIGURE 5d
FIGURE 5e

FIGURE 5e
FIGURE 5f

FIGURE 5f
FIGURE 5g

FIGURE 5g
FIGURE 5h

FIGURE 5h
Gallery 5 Myscelia, Paulogramma, Pyrrhogyra, Smyrna and Temenis
FIGURE 5a - Orsis Bluewing Myscelia orsis male - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 5b - Orsis Bluewing Myscelia orsis male - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 5c - Orsis Bluewing Myscelia orsis female - PROCOSARA, PN San Rafael (Laura Tensen February 2008 - ECOSARA Biodiversity Database).
FIGURE 5d - False Callicore Paulogramma pyracmon - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 5e - Redring Pyrrhogyra neaerea arge - Encarnación, Departamento Itapúa (Paul Smith May 2007).
FIGURE 5f - Redring Pyrrhogyra neaerea arge - upper image Encarnación, Departamento Itapúa (Paul Smith June 2007); lower image Hotel Tirol, Departamento Itapúa (Regis Nossent July 2005).
FIGURE 5g - Blomfild´s Beauty Smyrna blomfildia - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 5h - Mocha Temenis laothoe - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).

Notice the similarity of Paulogramma to the Callicore butterflies in Gallery 1. It is especially similar to C.pygas.
Hamadryas epinome territorial defence sounds recorded PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
Gallery 4 Hamadryas
FIGURE 4a - Red Clicker Hamadryas amphinome - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 4b - White Clicker Hamadryas epinome - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 4c - White Clicker Hamadryas epinome - PROCOSARA, PN San Rafael (Paul Smith upperwing March 2007, underwing June 2007 - ECOSARA Biodiversity Database).
FIGURE 4d - Grey Clicker Hamadryas februa - Encarnación, Departamento Itapúa (Paul Smith June 2007).
FIGURE 4e - Red-spotted Clicker Hamadryas feronia - Sanctuario de Itacua, Departamento Itapúa (Paul Smith September 2007).

Hamadryas butterflies are known as "Clickers" because of their ability to make clicking noises with the wings, perhaps as a territorial marker (Kim Garwood pers. comm.).
Click the link below to hear the noise made by the "White Clicker" Hamadryas epinome Figs 4b-c.
FIGURE 4a

FIGURE 4a
FIGURE 4b

FIGURE 4b
FIGURE 4c

FIGURE 4c
FIGURE 4d

FIGURE 4d
FIGURE 4e

FIGURE 4e