FIGURE 8a

FIGURE 8a
FIGURE 8b

FIGURE 8b
FIGURE 8c

FIGURE 8c
FIGURE 8d

FIGURE 8d
FIGURE 8e

FIGURE 8e
FIGURE 8f

FIGURE 8f
FIGURE 8g

FIGURE 8g
FIGURE 8h

FIGURE 8h
FIGURE 8i

FIGURE 8i
Thanks to Ezequiel Nuñez-Bustos and Richard Lindstrom for their assistance with skipper identification.
Designed by Paul Smith 2006. This website is copyrighted by law. Material contained herewith may not be used without the prior written permission of FAUNA Paraguay. Photographs on this web-site were taken by Paul Smith, Hemme Batjes, Regis Nossent,
Alberto Esquivel, Arne Lesterhuis, José Luis Cartes, Rebecca Zarza, and Hugo del Castillo and are used with their permission.
FIGURE 1a

FIGURE 1a
FIGURE 1b

FIGURE 1b
FIGURE 1c

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FIGURE 1d

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FIGURE 1f

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FIGURE 1g

FIGURE 1g
Gallery 1 Achylodes, Anisochoria and Anigonus
FIGURE 1a - Giant Sicklewing Achlyodes busirus - Encarnación, Departamento Itapúa (Paul Smith June 2007).
FIGURE 1b - Common Sicklewing Achylodes mithridates male - Encarnación, Departamento Itapúa (Paul Smith June 2007).
FIGURE 1c - Common Sicklewing Achylodes mithridates female - PROCOSARA, PN San Rafael (Moriz Stiefel April 2008 - ECOSARA Biodiversity Database).
FIGURE 1d - Anisochoria pedaliodina extincta - Estancia Kanguery, PN San Rafael (Paul Smith November 2007).
FIGURE 1e - Cut-wing Skipper Anisochoria sublimbata - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 1f - Antigonus erosus - PROCOSARA, PN San Rafael (Paul Smith March 2008 - ECOSARA Biodiversity Database).
FIGURE 1g - Antigonus nearchus - PROCOSARA, PN San Rafael (Paul Smith November 2007 - ECOSARA Biodiversity Database). 
SUBFAMILY PYRGINAE - FLAT-WINGED SKIPPERS

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Gallery 4 Dyscophellus, Ebrietas, Epargyreus, Gorgythion, Grais and Helias
FIGURE 4a- Dyscophellus erythras female - Hotel Tirol, Departamento Itapúa (Regis Nossent June 2005).
FIGURE 4b - Black-patched Duskywing Ebrietas anacreon  - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 4c - Colossus Skipper Epargyreus clavicornis - PROCOSARA, PN San Rafael (Paul Smith January 2008 - ECOSARA Biodiversity Database).
FIGURE 4d - Gorgythion begga - PROCOSARA, PN San Rafael (Jeni Oborn June 2008 - ECOSARA Biodiversity Database).
FIGURE 4e - Gorgythion beggina - Encarnación, Departamento Itapúa (Paul Smith April 2008).
FIGURE 4f - Grais stigmaticus - PROCOSARA, PN San Rafael (Laura Tensen March 2008 - ECOSARA Biodiversity Database).
FIGURE 4g - Helias phalaenoides - PROCOSARA, PN San Rafael (upper Moriz Stiefel, lower Paul Smith May 2008 - ECOSARA Biodiversity Database).
Gallery 8 Pythonides, Quadrus, Sostrata, Staphylus, Timochares and Trina
FIGURE 8a - Blue Lance Skipper Pythonides lancea - PROCOSARA, PN San Rafael (David Gill February 2008 - ECOSARA Biodiversity Database).
FIGURE 8b - Ceres´ Skipper Quadrus cerialis - PROCOSARA, PN San Rafael (Sylvia Qu May 2008 - ECOSARA Biodiversity Database).
FIGURE 8c - Lavender Skipper Sostrata bifasciata - PROCOSARA, PN San Rafael (Paul Smith November 2007 - ECOSARA Biodiversity Database).
FIGURE 8d - Sostrata cronion - PROCOSARA, PN San Rafael (Sylvia Qu May 2008 - ECOSARA Biodiversity Database).
FIGURE 8e - Staphylus ascalon - PROCOSARA, PN San Rafael (Paul Smith November 2007 - ECOSARA Biodiversity Database).
FIGURE 8f - Three-banded Skipper Timochares trifasciata - PROCOSARA, PN San Rafael (upper Paul Smith March 2007 lower Alberto Esquivel December 2007 - ECOSARA Biodiversity Database).
FIGURE 8g - Geometric Skipper Trina geometrina - PROCOSARA, PN San Rafael (Paul Smith February 2008 - ECOSARA Biodiversity Database).
FIGURE 8h - Geometric Skipper Trina geometrina - Sanctuario de Itacua, Departamento Itapúa (Paul Smith September 2007).
FIGURE 8i - Typhedanus undulatus - PROCOSARA, PN San Rafael (Paul Smith May 2008 - ECOSARA Biodiversity Database).
FIGURE 4a

FIGURE 4a
FIGURE 4b

FIGURE 4b
FIGURE 4c

FIGURE 4c
FIGURE 4d

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FIGURE 4e

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FIGURE 4f

FIGURE 4f
FIGURE 4g

FIGURE 4g
FIGURE 6a

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FIGURE 6c

FIGURE 6c
FIGURE 6d

FIGURE 6d
FIGURE 6e

FIGURE 6e
FIGURE 6f

FIGURE 6f
Gallery 6 Melanion, Narcosius, Nascus, Oechydrus,  Polygonus and Proteides and Pyrgus
FIGURE 6a White Shield Skipper Melanion leucaspis - PROCOSARA, PN San Rafael (Moriz Stiefel April 2008 - ECOSARA Biodiversity Database).
FIGURE 6b - Narcosius parisi - Hotel Tirol, Departamento Itapúa (Paul Smith September 2007).
FIGURE 6c - Common Scarlet-Eye Nascus phocus female - Encarnaciòn, Departamento Itapúa (Paul Smith upper October 2006 lower April 2008).
FIGURE 6d - Oechydrus chersis - Hotel Tirol, Departamento Itapúa (Paul Smith September 2007).
FIGURE 6e - Polygonus leo - Estancia Kanguery, PN San Rafael (Paul Smith November 2007 - ECOSARA Biodiversity Database).
FIGURE 6f - Mercurial Skipper Proteides mercurius - Encarnación, Departamento Itapúa (Paul Smith September 2007)
FIGURE 2a

FIGURE 2a
FIGURE 2b

FIGURE 2b
FIGURE 2c

FIGURE 2c
FIGURE 2d

FIGURE 2d
FIGURE 2e

FIGURE 2e
FIGURE 2f

FIGURE 2f
Gallery 9 Urbanus
FIGURE 9a - Lilac-banded Long-tailed Skipper Urbanus dorantes - PROCOSARA, PN San Rafael (Paul Smith May 2008 - ECOSARA Biodiversity Database).
FIGURE 9b - Urbanus doryssus - PROCOSARA, PN San Rafael (Jeni Oborn June 2008 - ECOSARA Biodiversity Database).
FIGURE 9c - Urbanus esta - PROCOSARA, PN San Rafael (Moriz Stiefel April 2008 - ECOSARA Biodiversity Database).
FIGURE 9d - Urbanus procne - Encarnación, Departamento Itapúa (Paul Smith May 2007).
FIGURE 9e - Urbanus procne - Encarnación, Departamento Itapúa (Paul Smith June 2007).
FIGURE 9f - Urbanus pronta - PROCOSARA, PN San Rafael (Nick Simms February 2008 - ECOSARA Biodiversity Database).
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FIGURE 9a
FIGURE 9b

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FIGURE 9c

FIGURE 9c
FIGURE 9d

FIGURE 9d
FIGURE 9e

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FIGURE 9f

FIGURE 9f
FIGURE 3a

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FIGURE 3c

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FIGURE 3d

FIGURE 3d
FIGURE 3e

FIGURE 3e
Gallery 2 Astraptes and Autochton
FIGURE 2a - Yellow-tailed Skipper Astraptes anaphus - PROCOSARA, PN San Rafael (Paul Smith February 2008 - ECOSARA Biodiversity Database). 
FIGURE 2b - Chalk-spotted Blue Skipper Astraptes creteus - Encarnación, Departamento Itapúa (Paul Smith April 2008).
FIGURE 2c - Astraptes elorus - PROCOSARA, PN San Rafael (Paul Smith March 2008 - ECOSARA Biodiversity Database).
FIGURE 2d - Astraptes fulgerator - PROCOSARA, PN San Rafael (Laura Tensen April 2008 - ECOSARA Biodiversity Database).
FIGURE 2e - Astraptes fulgor - PROCOSARA, PN San Rafael (Paul Smith May 2008 - ECOSARA Biodiversity Database).
FIGURE 2f - White-banded Skipper Autochton zarex - upper PROCOSARA, PN San Rafael (Paul Smith March 2008 - ECOSARA Biodiversity Database), lower Hotel Tirol, Departamento Itapúa (Paul Smith September 2007).
Gallery 3 Celaenorrhinus, Chioides, Chiomara, Codatractus,  and Cogia
FIGURE 3a - Obstinate Skipper Celaenorrhinus similis - PROCOSARA, PN San Rafael (upper Sylvia Qu June 2008; lower Moriz Stiefel May 2008 - ECOSARA Biodiversity Database). 
FIGURE 3b - White-striped Long-tailed Skipper Chioides catillus - PN Tte Enciso, Departamento Boquerón (Paul Smith July 2006).
FIGURE 3c - White-patched Skipper Chiomara asychis - PN Tte Enciso, Departamento Boquerón (Paul Smith July 2006).
FIGURE 3d - Gold-bordered Long-tailed Skipper Codatractus aminias - PROCOSARA, PN San Rafael (Paul Smith September 2007 - ECOSARA Biodiversity Database). 
FIGURE 3e - Cogia calchas - Encarnación, Departamento Itapúa (Paul Smith January 2008).
FIGURE 5a

FIGURE 5a
FIGURE 5b

FIGURE 5b
FIGURE 5c

FIGURE 5c
FIGURE 5d

FIGURE 5d
FIGURE 5e

FIGURE 5e
FIGURE 5f

FIGURE 5f
Gallery 5 Heliopetes and Mylon
FIGURE 5a - Alana´s White Skipper Heliopetes alana male? - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 5b - Alana´s White Skipper Heliopetes alana female? - Arroyo Mboi Kae, Departamento Itapúa (Regis Nossent July 2005).
FIGURE 5c - Heliopetes arsalte - Encarnaciòn, Departamento Itapúa (Paul Smith August 2007).
FIGURE 5d - Heliopetes libra female - PROCOSARA, PN San Rafael (Paul Smith February 2008 - ECOSARA Biodiversity Database).
FIGURE 5e - Heliopetes omrina - PROCOSARA, PN San Rafael (Paul Smith January 2008 - ECOSARA Biodiversity Database).
FIGURE 5f - Mylon maimon (=M.menippus) - PROCOSARA, PN San Rafael (Paul Smith upper March 2007; lower November 2007 - ECOSARA Biodiversity Database).
FIGURE 7a

FIGURE 7a
FIGURE 7b

FIGURE 7b
FIGURE 7c

FIGURE 7c
FIGURE 7d

FIGURE 7d
FIGURE 7e

FIGURE 7e
Gallery 7 Pyrgus
FIGURE 7a - Common Checkered Skipper Pyrgus orcynoides - PROCOSARA, PN San Rafael (Paul Smith March 2007 - ECOSARA Biodiversity Database).
FIGURE 7b - Common Checkered Skipper Pyrgus orcynoides - PROCOSARA, PN San Rafael (Martijn Brouwer May 2007 - ECOSARA Biodiversity Database).
FIGURE 7c - Pluto´s Checkered Skipper Pyrgus orcus male - PROCOSARA, PN San Rafael (upper image Martijn Brouwer May 2007, lower image Paul Smith November 2007 - ECOSARA Biodiversity Database).
FIGURE 7d - Pluto´s Checkered Skipper Pyrgus orcus female - PROCOSARA, PN San Rafael (Paul Smith November 2007 - ECOSARA Biodiversity Database).
FIGURE 7e - Pluto´s Checkered Skipper Pyrgus orcus mating pair - PROCOSARA, PN San Rafael (Paul Smith May 2008 - ECOSARA Biodiversity Database).
FIGURE 10a

FIGURE 10a
FIGURE 10b

FIGURE 10b
FIGURE 10c

FIGURE 10c
FIGURE 10d

FIGURE 10d
FIGURE 10e

FIGURE 10e
FIGURE 10f

FIGURE 10f
Gallery 10 Urbanus cont. and Xenophanes
FIGURE 10a - Turquoise Long-tailed Skipper Urbanus sp. - upper photo Arroyo Mboi Kae, Departamento Itapúa (Regis Nossent July 2005), lower photo Encarnación, Departamento Itapúa (Paul Smith May 2007).
FIGURE 10b - Plain Long-tailed Skipper Urbanus simplicus - Encarnación, Departamento Itapúa (Paul Smith May 2007).
FIGURE 10c - Lined Long-tailed Skipper Urbanus teleus - Arroyo Mboi Kae, Departamento Itapúa (Regis Nossent July 2005).
FIGURE 10d - Urbanus zagorus - PROCOSARA, PN San Rafael (Paul Smith November 2007 - ECOSARA Biodiversity Database).
FIGURE 10e - Glossy-winged Skipper Xenophanes tryxus - PROCOSARA, PN San Rafael (Martijn Brouwer May 2007 - ECOSARA Biodiversity Database).
FIGURE 10f - Glossy-winged Skipper Xenophanes tryxus - Hotel Tirol, Departamento Itapúa (Paul Smith September 2007).