FAUNA Paraguay ONLINE HANDBOOK OF PARAGUAYAN COLEOPTERA USERS GUIDE
Designed by Paul Smith 2006. This website is copyrighted by law. Material contained herewith may not be used without the prior written permission of FAUNA Paraguay. Photographs on this web-site were taken by Paul Smith, Hemme Batjes, Regis Nossent,
Alberto Esquivel, Arne Lesterhuis, Josť Luis Cartes, Rebecca Zarza and Hugo del Castillo and are used with their permission.
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Part of the ambitious FAUNA Paraguay Handbook of Paraguayan fauna project, the Online Coleoptera Handbook aims to give detailled information on the biology and identification of Paraguayan beetles in a readable, non-technical, style that will appeal to the interested amateur as much as the professional.
The species accounts are accessed via the links in the the familial and subfamilial image galleries
. Below is an explanation of how to use the species accounts.

Scientific name (Descriptor and date) Image Gallery (links to the image gallery of the taxonomic group to which the species belongs). The presence of rounded parantheses means that the species name has changed since the original description. If they are absent the species was described with the name it retains today.
TAX: Taxonomic issues and taxonomic history of the species.
ENG:
English name where one exists
ESP:
Spanish common name in general usage in Paraguay or Argentina where it exists.
DES: General description of the main characteristics of the species. The description omits general characters of shape and colouration that are common to the family and covered in the familial and subfamilial sections, concentrating only on characters that define the species.
MMT: The following measurements are given: TL - Total body length; EL - Length of the elytra or wing cases; WI - Width of the animal at its widest point;  n=number of measured specimens, Measurements are given in millimetres with the mean first followed by the range.
SSP:
Similar species. A paragraph distinguishing the species in question from its confusion species. Confusion species are linked in the text to allow quick cross reference.

ABU: Abundance and seasonality.
DIS: Distribution both in Paraguay and on a global scale.
HAB: Habitat preferences.
REP: Information on reproduction.
Citable Reference: This is the reference to be cited should the account be referred to in any other work.
Last Updated: Being an online handbook new material can be added as and when it becomes available. For reference purposes the date that the page was last updated is given here.

Images referred to in the above text are included here.